Pnas.org - SEO Checker

Overview of the SEO Check
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61% 
Page quality
42% 
Page structure
27% 
Link structure
25% 
Server
95% 
External factors
100% 
SEO Score
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2.13 s
File size
129.30 kB
Words
1965
Media files
24
Number of links
284 internal / 16 external

Task list of SEO Improvements

Meta specifications

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(Critically important)
PNAS
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Meta description
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Proceedings of the National Academy of Sciences (PNAS), a peer reviewed journal of the National Academy of Sciences (NAS) - an authoritative source of high-impact, original research that broadly spans the biological, physical, and social sciences.
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There are no problems in accessing the website.
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There is a valid canonical link specified.
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Language detected in text: en
Language defined in HTML: en
Server location: United States of America
The following language is defined by HTML: en
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og:titleProceedings of the National Academy of Sciences
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og:site_namePNAS
og:descriptionProceedings of the National Academy of Sciences (PNAS), a peer reviewed journal of the National Academy of Sciences (NAS) - an authoritative source of high-impact, original research that broadly spans the biological, physical, and social sciences.
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Content
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Words from the H1 heading are not used in the page content.
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  • Duplicate: This is an addendum toComplex viscosity of helical and doubly helical ...
The average number of words per sentence of 30 words is high.
This page contains 1965 words. That's ok.
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The usage of strong and bold tags is perfect. We recommend the use of up to 39 tags for this page.
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This website uses HTTPS to protect privacy and integrity of the exchanged data.
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Media list

URLAlt attributeTitle
...Logos/header-logo/logo-1624644560537.svgPNAS LogoPNAS logo
...header-logo/logo-light-1641218818190.svgPNAS LogoPNAS logo
...Logos/header-logo/logo-1624644560537.svgPNAS
...s/icons/twitter-x-gray-1705527808743.svgNo alt attribute provided
.../icons/researcher-gray-1705530250737.svgNo alt attribute provided
...tml/home-slider/slide0-1715698724823.jpgNo alt attribute provided
...tml/home-slider/slide1-1715698725740.jpgPictured are a caterpillar (bottom) and a biomimetic soft origami robot (top). Image credit: Glaucio H. Paulino.
...tml/home-slider/slide2-1715094611920.jpgRed wood ant carrying a pine needle.
...tml/home-slider/slide3-1701188922663.jpgIceberg
...9e/pnas.2024.121.issue-20.largecover.pngThe cover image for the current issue
...home/about-section-img-1624574253010.pngabout section
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...9e/pnas.2024.121.issue-20.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 20
...79/pnas.2024.121.issue-19.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 19
...f3/pnas.2024.121.issue-18.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 18
...c6/pnas.2024.121.issue-17.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 17
...dd/pnas.2024.121.issue-16.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 16
...c0/pnas.2024.121.issue-15.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 15
...e3/pnas.2024.121.issue-14.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 14
...f6/pnas.2024.121.issue-13.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 13
...f1/pnas.2024.121.issue-12.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 12
...gos/footer/logo-footer-1624408444100.svgPNAS logo

Page structure

H1 heading
(Critically important)
PNAS announces new website services! Dive deeper into the latest digital improvements in our Employee Spotlight blog series.
Too many H1 headings
The H1 heading is too long.
Headings
(Important)
Some headings occur twice on the page.
There are 63 headings on the page. The amount of headings should be in a more proper relation to the amount of text.

Heading structure

Heading levelContent
H1 PNAS announces new website services! Dive deeper into the latest digital improvements in our Employee Spotlight blog series.
H1 May 14, 2024; vol. 121 no. 20
H1 Latest in Applied Biological Sciences: Ants’ jaws could inspire the design of tools to manipulate objects in confined spaces.
H1 The world’s glaciers are melting. But how do researchers know exactly how much ice is disappearing and at what rate?
H2 Request Username
H2 Create a new account
H2 Login
H2 Change Password
H2 Current Issue
H2 PNAS is one of the world's most-cited and comprehensive multidisciplinary scientific journals.
H2 In the News
H2 Science Sessions podcasts
H2 Front Matter
H2 Featured QnAs
H2 Recent Issues
H2 Submit to PNAS
H3 In This Issue
H3 Modeling direct air carbon capture and storage in a 1.5 °C climate future using historical analogs
H3 Landscape fragmentation overturns classical metapopulation thinking
H3 Functional constraints on the number and shape of flight feathers
H3 A submerged Stone Age hunting architecture from the Western Baltic Sea
H3 Genomes, fossils, and the concurrent rise of modern birds and flowering plants in the Late Cretaceous
H3 Profile of Wanderley de Souza
H3 Bodily maps of emotions
H3 Oxytocin-enforced norm compliance reduces xenophobic outgroup rejection
H3 Neural network retuning and neural predictors of learning success associated with cello training
H3 Adapting to poor air quality
H3 Measuring Poverty
H3 How a small fish makes big sounds
H3 Making evidence go further: Advancing synergy between agent-based modeling and randomized control trials
H3 Community Review Boards offer a path to research equity
H3 “Net zero” may need a rethink to keep climate targets within reach
H3 The automated lab of tomorrow
H3 Species-level, digitized wildlife trade data are essential for achieving biodiversity targets
H3 We need a solid scientific basis for nature-based climate solutions in the United States
H3 To be policy-relevant, future climate research must include the noncontiguous United States
H3 Fire in the desert
H3 QnAs with Katherine Freese
H3 QnAs with Miguel A. Garcia-Garibay
H3 QnAs with Eugenia Russinova
H3 Sign up for thePNAS Highlights newsletter
H4 Featured Topics
H4 Articles By Topic
H4 Featured Topics Duplicate text
H4 Articles By Topic Duplicate text
H4 Featured Topic
H4 Articles By Topic Duplicate text
H4 PNAS UPDATES
H4 Announcing New Website Services: Expanded Author Services, Updates to the Article PDF, and Article-Podcast Links
H4 2023 Cozzarelli Prize Recipients
H4 PNAS Nexus Expands Its License Options
H4 PNAS Announces New Guidelines for the Retention of Raw Data
H4 Selected highlights of 2023
H5 Congrats!
H5 Vol. 121No. 20
H5 Vol. 121No. 19
H5 Vol. 121No. 18
H5 Vol. 121No. 17
H5 Vol. 121No. 16
H5 Vol. 121No. 15
H5 Vol. 121No. 14
H5 Vol. 121No. 13
H5 Vol. 121No. 12
Some internal link anchor texts are too long.
Some internal links have dynamic parameters. All internal URLs, which are not marked as nofollow, should not contain dynamic parameters.
Some anchor texts are used more than once.
There are 3 links with a trivial anchor text.
The number of internal links is ok.
There are 16 external links on this page.
LinkAttributesAnchor text
https://www.pnas.org/IMG-ALT PNAS Logo
https://www.pnas.org/Text duplicate IMG-ALT PNAS Logo
/toc/pnas/currentCurrent Issue
https://www.pnas.org/latestLatest Articles
/about/special-featuresSpecial Features
https://www.pnas.org/loi/pnasList of Issues
https://academic.oup.com/pnasn...External Subdomain PNAS Nexus
https://www.pnas.org/front-matterFront Matter
/author-centerInformation for Authors
/author-center/editorial-and-j...Editorial and Journal Policies
/author-center/submitting-your...Submission Procedures
/author-center/publication-cha...Publication Charges
https://www.pnas.org/Anchor Physical Sciences
https://www.pnas.org/Anchor Social Sciences
https://www.pnas.org/Anchor Biological Sciences
https://www.pnas.org/topic/physPhysics
https://www.pnas.org/topic/chemChemistry
/sustainability-scienceSustainability Science
/topic/app-mathApplied Mathematics
/topic/app-physApplied Physical Sciences
https://www.pnas.org/topic/astroAstronomy
/biophysics-and-computational-...Biophysics and Computational Biology
/topic/comp-sciComputer Sciences
/topic/earth-sciEarth, Atmospheric, and Planetary Sciences
https://www.pnas.org/topic/engEngineering
/environmental-sciencesEnvironmental Sciences
https://www.pnas.org/topic/mathMathematics
https://www.pnas.org/topic/statStatistics
https://www.pnas.org/anthropologyAnthropology
/sustainability-scienceText duplicate Sustainability Science
https://www.pnas.org/topic/demoDemography
/topic/econ-sciEconomic Sciences
/environmental-sciencesText duplicate Environmental Sciences
/topic/pol-sciPolitical Sciences
/psychological-and-cognitive-s...Psychological and Cognitive Sciences
https://www.pnas.org/anthropologyText duplicate Anthropology
/sustainability-scienceText duplicate Sustainability Science
https://www.pnas.org/topic/ag-sciAgricultural Sciences
/topic/app-bioApplied Biological Sciences
/topic/biochemBiochemistry
/biophysics-and-computational-...Text duplicate Biophysics and Computational Biology
/topic/cell-bioCell Biology
/topic/dev-bioDevelopmental Biology
https://www.pnas.org/topic/ecoEcology
/environmental-sciencesText duplicate Environmental Sciences
/topic/evolutionEvolution
/topic/geneticsGenetics
https://www.pnas.org/topic/immunImmunology and Inflammation
/topic/med-sciMedical Sciences
/topic/microbioMicrobiology
https://www.pnas.org/topic/neuroNeuroscience
https://www.pnas.org/topic/pharmPharmacology
https://www.pnas.org/topic/physioPhysiology
/topic/plant-bioPlant Biology
/topic/pop-bioPopulation Biology
/psychological-and-cognitive-s...Text duplicate Psychological and Cognitive Sciences
/topic/sys-bioSystems Biology
/topic/covid-19Covid-19
/action/showCart?FlowID=10 Cart
A-TITLE Show shopping cart
/action/showLoginSign in
/action/registrationRegister
/action/showLogin?redirectUri=/Individual Login
/action/ssostart?redirectUri=/Institutional Login
https://www.pnascentral.org/cg...External Subdomain Submit
https://www.pnas.org/Anchor Anywhere
https://www.pnas.org/Anchor Citation
https://www.pnas.org/Anchor Text duplicate PNAS Nexus
https://www.pnas.org/Anchor Proceedings of the National Academy of Sciences
/search/advancedAdvanced Search
/action/doSearch?AllField=covi...Text duplicate Covid-19
/action/doSearch?AllField=arti...Artificial Intelligence
/action/doSearch?AllField=mpoxMpox
/action/doSearch?AllField=clim...Climate Change
/action/doSearch?AllField=gun+...Gun Violence
https://www.pnas.org/IMG-ALT PNAS
https://www.facebook.com/pages...External Subdomain A-TITLE Facebook
https://twitter.com/PNASNewsExternal A-TITLE Twitter
https://www.youtube.com/channe...External Subdomain A-TITLE Youtube
https://www.linkedin.com/showc...New window External Subdomain A-TITLE LinkedIn
/action/showPreferences?menuTa...A-TITLE Mail
https://www.pnas.org/about/rssA-TITLE RSS feeds
https://www.researcher-app.com...External Subdomain A-TITLE Researcher App
/about/staff-directorySubdomain Contact
/about/site-mapSubdomain Site Map
https://www.pnas.org/about/termsSubdomain Terms & Privacy Policy
/about/accessibilitySubdomain Accessibility
https://www.pnas.org/Anchor Skip slideshow
/post/update/new-website-servi...PNAS announces new website services! Dive deeper into the latest digital improvements in our Employee Spotlight blog series. Read now.
/toc/pnas/121/20May 14, 2024; vol. 121 no. 20 Volume 121, Issue 20: View articles in the latest issue of PNAS.
IMG-ALT Pictured are a caterpillar (bottom) and a biomimetic soft origami robot (top). Image credit: Glaucio H. Paulino.
https://doi.org/10.1073/pnas.2...External Latest in Applied Biological Sciences: Ants’ jaws could inspire the design of tools to manipulate objects in confined spaces. Click to read more. Image credi...
IMG-ALT Red wood ant carrying a pine needle.
/post/update/measure-massive-meltThe world’s glaciers are melting. But how do researchers know exactly how much ice is disappearing and at what rate? Click to watch the Featured Video.
IMG-ALT Iceberg
/action/showPreferences?menuTa...sign up for alerts
/author-centerAuthor center
https://www.pnascentral.org/cg...External Subdomain submit
/action/showLoginReturning user
/help/cookie-policyFind out more
/help/cookie-policyCONTINUE
/action/requestResetPasswordTrivial anchor text
here
/action/registrationText duplicate Trivial anchor text
here
/action/ssostart?redirectUri=/Text duplicate Institutional Login
/toc/pnas/121/20Text duplicate Current Issue
/toc/pnas/currentIMG-ALT The cover image for the current issue
A-TITLE View the current issue table of contents
/topic/type/twipThis Week in PNAS
/doi/10.1073/iti2024121In This Issue
A-TITLE In This Issue
/topic/type/research-articleResearch Article
/doi/10.1073/pnas.2215679121Modeling direct air carbon capture and storage in a 1.5 °C climate future using historical analogs
A-TITLE Modeling direct air carbon capture and storage in a 1.5 °C climate future using historical analogs
/author/Edwards,+Morgan+RMorgan R. Edwards
A-TITLE Morgan R. Edwards
/author/Thomas,+Zachary+HZachary H. Thomas
A-TITLE Zachary H. Thomas
/author/Nemet,+Gregory+FGregory F. Nemet
A-TITLE Gregory F. Nemet
/author/Rathod,+SagarSagar Rathod
A-TITLE Sagar Rathod
/author/Greene,+JennaJenna Greene
A-TITLE Jenna Greene
/author/Surana,+KavitaKavita Surana
A-TITLE Kavita Surana
/author/Kennedy,+Kathleen+MKathleen M. Kennedy
A-TITLE Kathleen M. Kennedy
/author/Fuhrman,+JayJay Fuhrman
A-TITLE Jay Fuhrman
/author/McJeon,+Haewon+CHaewon C. McJeon
A-TITLE Haewon C. McJeon
/topic/type/research-articleText duplicate Research Article
/doi/10.1073/pnas.2303846121Landscape fragmentation overturns classical metapopulation thinking
A-TITLE Landscape fragmentation overturns classical metapopulation thinking
/author/Tao,+YunYun Tao
A-TITLE Yun Tao
/author/Hastings,+AlanAlan Hastings
A-TITLE Alan Hastings
/author/Lafferty,+Kevin+DKevin D. Lafferty
A-TITLE Kevin D. Lafferty
/author/Hanski,+IlkkaIlkka Hanski
A-TITLE Ilkka Hanski
/author/Ovaskainen,+OtsoOtso Ovaskainen
A-TITLE Otso Ovaskainen
/toc/pnas/currentVIEW FULL TABLE OF CONTENTS
https://www.pnas.org/aboutTrivial anchor text
Read More
/topic/type/research-articleText duplicate Research Article
/doi/10.1073/pnas.2306639121Functional constraints on the number and shape of flight feathers
A-TITLE Functional constraints on the number and shape of flight feathers
/author/Kiat,+YosefYosef Kiat
A-TITLE Yosef Kiat
/author/O'Connor,+Jingmai+KJingmai K. O’Connor
A-TITLE Jingmai K. O’Connor
/topic/type/research-articleText duplicate Research Article
/doi/10.1073/pnas.2312008121A submerged Stone Age hunting architecture from the Western Baltic Sea
A-TITLE A submerged Stone Age hunting architecture from the Western Baltic Sea
/author/Geersen,+JacobJacob Geersen
A-TITLE Jacob Geersen
/author/Bradtmöller,+MarcelMarcel Bradtmöller
A-TITLE Marcel Bradtmöller
/author/Schneider+von+Deimling...Jens Schneider von Deimling
A-TITLE Jens Schneider von Deimling
/author/Feldens,+PeterPeter Feldens
A-TITLE Peter Feldens
/author/Auer,+JensJens Auer
A-TITLE Jens Auer
/author/Held,+PhilippPhilipp Held
A-TITLE Philipp Held
/author/Lohrberg,+ArneArne Lohrberg
A-TITLE Arne Lohrberg
/author/Supka,+RuthRuth Supka
A-TITLE Ruth Supka
/author/Hoffmann,+Jasper+Justu...Jasper Justus Lutz Hoffmann
A-TITLE Jasper Justus Lutz Hoffmann
/author/Eriksen,+Berit+ValentinBerit Valentin Eriksen
A-TITLE Berit Valentin Eriksen
/author/Rabbel,+WolfgangWolfgang Rabbel
A-TITLE Wolfgang Rabbel
/author/Karlsen,+Hans-JörgHans-Jörg Karlsen
A-TITLE Hans-Jörg Karlsen
/author/Krastel,+SebastianSebastian Krastel
A-TITLE Sebastian Krastel
/author/Brandt,+DavidDavid Brandt
A-TITLE David Brandt
/author/Heuskin,+DavidDavid Heuskin
A-TITLE David Heuskin
/author/Lübke,+HaraldHarald Lübke
A-TITLE Harald Lübke
/topic/type/research-articleText duplicate Research Article
/doi/10.1073/pnas.2319696121Genomes, fossils, and the concurrent rise of modern birds and flowering plants in the Late Cretaceous
A-TITLE Genomes, fossils, and the concurrent rise of modern birds and flowering plants in the Late Cretaceous
/author/Wu,+ShaoyuanShaoyuan Wu
A-TITLE Shaoyuan Wu
/author/Rheindt,+Frank+EFrank E. Rheindt
A-TITLE Frank E. Rheindt
/author/Zhang,+JinJin Zhang
A-TITLE Jin Zhang
/author/Wang,+JiajiaJiajia Wang
A-TITLE Jiajia Wang
/author/Zhang,+LeiLei Zhang
A-TITLE Lei Zhang
/author/Quan,+ChengCheng Quan
A-TITLE Cheng Quan
/author/Li,+ZhihengZhiheng Li
A-TITLE Zhiheng Li
/author/Wang,+MinMin Wang
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/author/Wu,+FeixiangFeixiang Wu
A-TITLE Feixiang Wu
/author/Qu,+YanhuaYanhua Qu
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/author/Edwards,+Scott+VScott V. Edwards
A-TITLE Scott V. Edwards
/author/Zhou,+ZhongheZhonghe Zhou
A-TITLE Zhonghe Zhou
/author/Liu,+LiangLiang Liu
A-TITLE Liang Liu
/topic/type/profileProfile
/doi/10.1073/pnas.2403579121Profile of Wanderley de Souza
A-TITLE Profile of Wanderley de Souza
/author/Viegas,+JenniferJennifer Viegas
A-TITLE Jennifer Viegas
/about/pnas-in-the-newsView all news
/topic/type/research-articleText duplicate Research Article
/doi/full/10.1073/pnas.1321664111Bodily maps of emotions
A-TITLE Bodily maps of emotions
/author/Nummenmaa,+LauriLauri Nummenmaa
A-TITLE Lauri Nummenmaa
/author/Glerean,+EnricoEnrico Glerean
A-TITLE Enrico Glerean
/author/Hari,+RiittaRiitta Hari
A-TITLE Riitta Hari
/author/Hietanen,+Jari+KJari K. Hietanen
A-TITLE Jari K. Hietanen
/topic/type/research-articleText duplicate Research Article
/doi/full/10.1073/pnas.1705853114Oxytocin-enforced norm compliance reduces xenophobic outgroup rejection
A-TITLE Oxytocin-enforced norm compliance reduces xenophobic outgroup rejection
/author/Marsh,+NinaNina Marsh
A-TITLE Nina Marsh
/author/Scheele,+DirkDirk Scheele
A-TITLE Dirk Scheele
/author/Feinstein,+Justin+SJustin S. Feinstein
A-TITLE Justin S. Feinstein
/author/Gerhardt,+HolgerHolger Gerhardt
A-TITLE Holger Gerhardt
/author/Strang,+SabrinaSabrina Strang
A-TITLE Sabrina Strang
/author/Maier,+WolfgangWolfgang Maier
A-TITLE Wolfgang Maier
/author/Hurlemann,+RenéRené Hurlemann
A-TITLE René Hurlemann
/topic/type/research-articleText duplicate Research Article
/doi/full/10.1073/pnas.1721414115Neural network retuning and neural predictors of learning success associated with cello training
A-TITLE Neural network retuning and neural predictors of learning success associated with cello training
/author/Wollman,+IndianaIndiana Wollman
A-TITLE Indiana Wollman
/author/Penhune,+VirginiaVirginia Penhune
A-TITLE Virginia Penhune
/author/Segado,+MelanieMelanie Segado
A-TITLE Melanie Segado
/author/Carpentier,+ThibautThibaut Carpentier
A-TITLE Thibaut Carpentier
/author/Zatorre,+Robert+JRobert J. Zatorre
A-TITLE Robert J. Zatorre
https://www.pnas.org/trendingView more
/doi/10.1073/pc.31102568play Adapting to poor air quality
IMG-ALT placeholder
/doi/10.1073/pc.30908323play Measuring Poverty
IMG-ALT placeholder
/doi/10.1073/pc.30695898play How a small fish makes big sounds
IMG-ALT placeholder
/about/science-sessions-podcastmore Science Sessions
/topic/type/opinionOpinion
/doi/10.1073/pnas.2314993121Making evidence go further: Advancing synergy between agent-based modeling and randomized control trials
A-TITLE Making evidence go further: Advancing synergy between agent-based modeling and randomized control trials
/author/Hammond,+Ross+ARoss A. Hammond
A-TITLE Ross A. Hammond
/author/Barkin,+ShariShari Barkin
A-TITLE Shari Barkin
/topic/type/opinionText duplicate Opinion
/doi/10.1073/pnas.2320334121Community Review Boards offer a path to research equity
A-TITLE Community Review Boards offer a path to research equity
/author/Sabet,+Cameron+JohnCameron John Sabet
A-TITLE Cameron John Sabet
/author/Bajaj,+Simar+SSimar S. Bajaj
A-TITLE Simar S. Bajaj
/author/Stanford,+Fatima+CodyFatima Cody Stanford
A-TITLE Fatima Cody Stanford
/topic/type/core-conCore Concepts
/doi/10.1073/pnas.2407160121“Net zero” may need a rethink to keep climate targets within reach
A-TITLE “Net zero” may need a rethink to keep climate targets within reach
/author/Battersby,+StephenStephen Battersby
A-TITLE Stephen Battersby
/topic/type/news-featNews Feature
/doi/10.1073/pnas.2406320121The automated lab of tomorrow
A-TITLE The automated lab of tomorrow
/author/Adam,+DavidDavid Adam
A-TITLE David Adam
/topic/type/opinionText duplicate Opinion
/doi/10.1073/pnas.2306869121Species-level, digitized wildlife trade data are essential for achieving biodiversity targets
A-TITLE Species-level, digitized wildlife trade data are essential for achieving biodiversity targets
/author/Tlusty,+Michael+FMichael F. Tlusty
A-TITLE Michael F. Tlusty
/author/Cassey,+PhillipPhillip Cassey
A-TITLE Phillip Cassey
/author/Rhyne,+Andrew+LAndrew L. Rhyne
A-TITLE Andrew L. Rhyne
/author/Omrow,+Delon+ADelon A. Omrow
A-TITLE Delon A. Omrow
/author/Stoett,+PeterPeter Stoett
A-TITLE Peter Stoett
/topic/type/opinionText duplicate Opinion
/doi/10.1073/pnas.2318505121We need a solid scientific basis for nature-based climate solutions in the United States
A-TITLE We need a solid scientific basis for nature-based climate solutions in the United States
/author/Novick,+Kimberly+AKimberly A. Novick
A-TITLE Kimberly A. Novick
/author/Keenan,+Trevor+FTrevor F. Keenan
A-TITLE Trevor F. Keenan
/author/Anderegg,+William+R+LWilliam R. L. Anderegg
A-TITLE William R. L. Anderegg
/author/Normile,+Caroline+PCaroline P. Normile
A-TITLE Caroline P. Normile
/author/Runkle,+Benjamin+R+KBenjamin R. K. Runkle
A-TITLE Benjamin R. K. Runkle
/author/Oldfield,+Emily+EEmily E. Oldfield
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