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61% 
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29% 
Link structure
25% 
Server
95% 
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100% 
SEO Score
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1.71 s
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133.10 kB
Words
1962
Media files
25
Number of links
302 internal / 16 external

Task list of SEO Improvements

Meta specifications

Title
(Critically important)
PNAS
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Meta description
(Critically important)
Proceedings of the National Academy of Sciences (PNAS), a peer reviewed journal of the National Academy of Sciences (NAS) - an authoritative source of high-impact, original research that broadly spans the biological, physical, and social sciences.
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There are no problems in accessing the website.
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There is a valid canonical link specified.
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Language detected in text: en
Language defined in HTML: en
Server location: United States of America
The following language is defined by HTML: en
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og:descriptionProceedings of the National Academy of Sciences (PNAS), a peer reviewed journal of the National Academy of Sciences (NAS) - an authoritative source of high-impact, original research that broadly spans the biological, physical, and social sciences.
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Words from the H1 heading are not used in the page content.
There are 21 text duplicates on this page:
  • Duplicate: This is an addendum toComplex viscosity of helical and doubly helical ...
The average number of words per sentence of 29.82 words is high.
This page contains 1962 words. That's ok.
24.1% of the text are stop words.
The page contains a listing, which indicates a good text layout.
31 paragraphs were found on this page.
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A viewport "width=device-width,initial-scale=1" is provided.
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This website uses HTTPS to protect privacy and integrity of the exchanged data.
All included files are also transferred via HTTPS.

Media list

URLAlt attributeTitle
...Logos/header-logo/logo-1624644560537.svgPNAS LogoPNAS logo
...header-logo/logo-light-1641218818190.svgPNAS LogoPNAS logo
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...s/icons/twitter-x-gray-1705527808743.svgNo alt attribute provided
.../icons/researcher-gray-1705530250737.svgNo alt attribute provided
...tml/home-slider/slide0-1723562634037.jpgNo alt attribute provided
...tml/home-slider/slide1-1723562634467.jpgPictured is a wafer-scale, flexible spectrometer array. Image credit: Chunyu You, Ningge Huang, and Yongfeng Mei.
...tml/home-slider/slide2-1723562634073.jpgXenophrys mangshanensis in the Nanling Mountains.
...tml/home-slider/slide3-1701188922663.jpgIceberg
...92/pnas.2024.121.issue-33.largecover.pngThe cover image for the current issue
...home/about-section-img-1624574253010.pngabout section
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...92/pnas.2024.121.issue-33.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 33
...02/pnas.2024.121.issue-32.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 32
...8e/pnas.2024.121.issue-31.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 31
...3d/pnas.2024.121.issue-30.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 30
...65/pnas.2024.121.issue-29.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 29
...ce/pnas.2024.121.issue-28.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 28
...77/pnas.2024.121.issue-27.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 27
...f0/pnas.2024.121.issue-26.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 26
...1c/pnas.2024.121.issue-25.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 25
...9e/pnas.2024.121.issue-24.largecover.pngThe cover image for PNAS Table of Contents Vol. 121 No. 24
...gos/footer/logo-footer-1624408444100.svgPNAS logo

Page structure

H1 heading
(Critically important)
Read PNAS Highlights—a collection of top scientific research, news, and profiles delivered to your inbox twice a month.
Too many H1 headings
Headings
(Important)
Some headings occur twice on the page.
There are 64 headings on the page. The amount of headings should be in a more proper relation to the amount of text.

Heading structure

Heading levelContent
H1 Read PNAS Highlights—a collection of top scientific research, news, and profiles delivered to your inbox twice a month.
H1 August 13, 2024; vol. 121 no. 33
H1 Latest in Ecology: Amphibian biodiversity hotspots in China
H1 The world’s glaciers are melting. But how do researchers know exactly how much ice is disappearing and at what rate?
H2 Request Username
H2 Create a new account
H2 Login
H2 Change Password
H2 Current Issue
H2 PNAS is one of the world's most-cited and comprehensive multidisciplinary scientific journals.
H2 In the News
H2 Science Sessions podcasts
H2 Front Matter
H2 Featured QnAs
H2 Recent Issues
H2 Submit to PNAS
H3 In This Issue
H3 Spatiotemporally controlled microvortices provide advanced microfluidic components
H3 Understanding the relationship between intergenerational mobility and community violence
H3 The thermoneutral zone in women takes an “arctic” shift compared to men
H3 Effect of skull morphology on fox snow diving
H3 Atmosphere teleconnections from abatement of China aerosol emissions exacerbate Northeast Pacific warm blob events
H3 QnAs with Spyros Artavanis-Tsakonas
H3 Liquid water in the Martian mid-crust
H3 A therapeutic small molecule enhances γ-oscillations and improves cognition/memory in Alzheimer’s disease model mice
H3 Bodily maps of emotions
H3 How redlining affects biodiversity
H3 Animal's eye view of the ocean
H3 Inequitable exposure to wildfire smoke
H3 Extended time, elevated expectations: The unappreciated downsides of pausing the tenure clock
H3 Synthetic data can aid the analysis of clinical outcomes: How much can it be trusted?
H3 Tiny and contentious “crystal clocks” in volcanic rocks could aid early warning for eruptions
H3 Researchers need to better address time-of-day as a critical biological variable
H3 Collaborative consortia can boost postdoctoral workforce development
H3 Sediment DNA can revolutionize archaeology—if it is used the right way
H3 Can ChatGPT help researchers understand how the human brain handles language?
H3 The Global Biodiversity Framework can be leveraged to better manage transboundary watersheds
H3 QnAs with Susan T. Lovett
H3 QnAs with Ulrich R. Christensen
H3 QnAs with Spyros Artavanis-Tsakonas Duplicate text
H3 Sign up for thePNAS Highlights newsletter
H4 Featured Topics
H4 Articles By Topic
H4 Featured Topics Duplicate text
H4 Articles By Topic Duplicate text
H4 Featured Topic
H4 Articles By Topic Duplicate text
H4 PNAS UPDATES
H4 PNAS Special Feature Blog: Physics Meets Machine Learning
H4 Announcing New Website Services: Expanded Author Services, Updates to the Article PDF, and Article-Podcast Links
H4 2023 Cozzarelli Prize Recipients
H4 PNAS Nexus Expands Its License Options
H4 PNAS Announces New Guidelines for the Retention of Raw Data
H5 Congrats!
H5 Vol. 121No. 33
H5 Vol. 121No. 32
H5 Vol. 121No. 31
H5 Vol. 121No. 30
H5 Vol. 121No. 29
H5 Vol. 121No. 28
H5 Vol. 121No. 27
H5 Vol. 121No. 26
H5 Vol. 121No. 25
H5 Vol. 121No. 24
Some internal link anchor texts are too long.
Some internal links have dynamic parameters. All internal URLs, which are not marked as nofollow, should not contain dynamic parameters.
Some anchor texts are used more than once.
There are 3 links with a trivial anchor text.
The number of internal links is ok.
There are 16 external links on this page.

Found links on this page

LinkAttributesAnchor text
https://www.pnas.org/IMG-ALT PNAS Logo
https://www.pnas.org/Text duplicate IMG-ALT PNAS Logo
/toc/pnas/currentCurrent Issue
https://www.pnas.org/latestLatest Articles
/about/special-featuresSpecial Features
https://www.pnas.org/loi/pnasList of Issues
https://academic.oup.com/pnasn...External Subdomain PNAS Nexus
https://www.pnas.org/front-matterFront Matter
/author-centerInformation for Authors
/author-center/editorial-and-j...Editorial and Journal Policies
/author-center/submitting-your...Submission Procedures
/author-center/publication-cha...Publication Charges
https://www.pnas.org/Anchor Physical Sciences
https://www.pnas.org/Anchor Social Sciences
https://www.pnas.org/Anchor Biological Sciences
https://www.pnas.org/topic/physPhysics
https://www.pnas.org/topic/chemChemistry
/sustainability-scienceSustainability Science
/topic/app-mathApplied Mathematics
/topic/app-physApplied Physical Sciences
https://www.pnas.org/topic/astroAstronomy
/biophysics-and-computational-...Biophysics and Computational Biology
/topic/comp-sciComputer Sciences
/topic/earth-sciEarth, Atmospheric, and Planetary Sciences
https://www.pnas.org/topic/engEngineering
/environmental-sciencesEnvironmental Sciences
https://www.pnas.org/topic/mathMathematics
https://www.pnas.org/topic/statStatistics
https://www.pnas.org/anthropologyAnthropology
/sustainability-scienceText duplicate Sustainability Science
https://www.pnas.org/topic/demoDemography
/topic/econ-sciEconomic Sciences
/environmental-sciencesText duplicate Environmental Sciences
/topic/pol-sciPolitical Sciences
/psychological-and-cognitive-s...Psychological and Cognitive Sciences
https://www.pnas.org/anthropologyText duplicate Anthropology
/sustainability-scienceText duplicate Sustainability Science
https://www.pnas.org/topic/ag-sciAgricultural Sciences
/topic/app-bioApplied Biological Sciences
/topic/biochemBiochemistry
/biophysics-and-computational-...Text duplicate Biophysics and Computational Biology
/topic/cell-bioCell Biology
/topic/dev-bioDevelopmental Biology
https://www.pnas.org/topic/ecoEcology
/environmental-sciencesText duplicate Environmental Sciences
/topic/evolutionEvolution
/topic/geneticsGenetics
https://www.pnas.org/topic/immunImmunology and Inflammation
/topic/med-sciMedical Sciences
/topic/microbioMicrobiology
https://www.pnas.org/topic/neuroNeuroscience
https://www.pnas.org/topic/pharmPharmacology
https://www.pnas.org/topic/physioPhysiology
/topic/plant-bioPlant Biology
/topic/pop-bioPopulation Biology
/psychological-and-cognitive-s...Text duplicate Psychological and Cognitive Sciences
/topic/sys-bioSystems Biology
/topic/covid-19Covid-19
/action/showCart?FlowID=10 Cart
A-TITLE Show shopping cart
/action/showLoginSign in
/action/registrationRegister
/action/showLogin?redirectUri=/Individual Login
/action/ssostart?redirectUri=/Institutional Login
https://www.pnascentral.org/cg...External Subdomain Submit
https://www.pnas.org/Anchor Anywhere
https://www.pnas.org/Anchor Citation
https://www.pnas.org/Anchor Text duplicate PNAS Nexus
https://www.pnas.org/Anchor Proceedings of the National Academy of Sciences
/search/advancedAdvanced Search
/action/doSearch?AllField=covi...Text duplicate Covid-19
/action/doSearch?AllField=arti...Artificial Intelligence
/action/doSearch?AllField=mpoxMpox
/action/doSearch?AllField=clim...Climate Change
/action/doSearch?AllField=gun+...Gun Violence
https://www.pnas.org/IMG-ALT PNAS
https://www.facebook.com/pages...External Subdomain A-TITLE Facebook
https://twitter.com/PNASNewsExternal A-TITLE Twitter
https://www.youtube.com/channe...External Subdomain A-TITLE Youtube
https://www.linkedin.com/showc...New window External Subdomain A-TITLE LinkedIn
/action/showPreferences?menuTa...A-TITLE Mail
https://www.pnas.org/about/rssA-TITLE RSS feeds
https://www.researcher-app.com...External Subdomain A-TITLE Researcher App
/about/staff-directorySubdomain Contact
/about/site-mapSubdomain Site Map
https://www.pnas.org/about/termsSubdomain Terms & Privacy Policy
/about/accessibilitySubdomain Accessibility
https://www.pnas.org/Anchor Skip slideshow
http://eepurl.com/gymAfDExternal Read PNAS Highlights—a collection of top scientific research, news, and profiles delivered to your inbox twice a month. Sign up here.
/toc/pnas/121/33August 13, 2024; vol. 121 no. 33 Volume 121, Issue 33: View articles in the latest issue of PNAS.
IMG-ALT Pictured is a wafer-scale, flexible spectrometer array. Image credit: Chunyu You, Ningge Huang, and Yongfeng Mei.
https://doi.org/10.1073/pnas.2...External Latest in Ecology: Amphibian biodiversity hotspots in China Click to read more. Image credit: Han Wan.
IMG-ALT Xenophrys mangshanensis in the Nanling Mountains.
/post/update/measure-massive-meltThe world’s glaciers are melting. But how do researchers know exactly how much ice is disappearing and at what rate? Click to watch the Featured Video.
IMG-ALT Iceberg
/action/showPreferences?menuTa...sign up for alerts
/author-centerAuthor center
https://www.pnascentral.org/cg...External Subdomain submit
/action/showLoginReturning user
/help/cookie-policyFind out more
/help/cookie-policyCONTINUE
/action/requestResetPasswordTrivial anchor text
here
/action/registrationText duplicate Trivial anchor text
here
/action/ssostart?redirectUri=/Text duplicate Institutional Login
/toc/pnas/121/33Text duplicate Current Issue
/toc/pnas/currentIMG-ALT The cover image for the current issue
A-TITLE View the current issue table of contents
/topic/type/twipThis Week in PNAS
/doi/10.1073/iti3324121In This Issue
A-TITLE In This Issue
/topic/type/research-articleResearch Article
/doi/10.1073/pnas.2306182121Spatiotemporally controlled microvortices provide advanced microfluidic components
A-TITLE Spatiotemporally controlled microvortices provide advanced microfluidic components
/author/Saito,+MakotoMakoto Saito
A-TITLE Makoto Saito
/author/Arai,+FumihitoFumihito Arai
A-TITLE Fumihito Arai
/author/Yamanishi,+YokoYoko Yamanishi
A-TITLE Yoko Yamanishi
/author/Sakuma,+ShinyaShinya Sakuma
A-TITLE Shinya Sakuma
/topic/type/research-articleText duplicate Research Article
/doi/10.1073/pnas.2309066121Understanding the relationship between intergenerational mobility and community violence
A-TITLE Understanding the relationship between intergenerational mobility and community violence
/author/Mann,+OliviaOlivia Mann
A-TITLE Olivia Mann
/author/Edin,+Kathryn+JKathryn J. Edin
A-TITLE Kathryn J. Edin
/author/Shaefer,+H+LukeH. Luke Shaefer
A-TITLE H. Luke Shaefer
/toc/pnas/currentVIEW FULL TABLE OF CONTENTS
https://www.pnas.org/aboutTrivial anchor text
Read More
/topic/type/research-articleText duplicate Research Article
/doi/10.1073/pnas.2311116121The thermoneutral zone in women takes an “arctic” shift compared to men
A-TITLE The thermoneutral zone in women takes an “arctic” shift compared to men
/author/Brychta,+Robert+JRobert J. Brychta
A-TITLE Robert J. Brychta
/author/McGehee,+SuzanneSuzanne McGehee
A-TITLE Suzanne McGehee
/author/Huang,+ShanShan Huang
A-TITLE Shan Huang
/author/Leitner,+Brooks+PBrooks P. Leitner
A-TITLE Brooks P. Leitner
/author/Duckworth,+Courtney+JCourtney J. Duckworth
A-TITLE Courtney J. Duckworth
/author/Fletcher,+Laura+ALaura A. Fletcher
A-TITLE Laura A. Fletcher
/author/Kim,+KatherineKatherine Kim
A-TITLE Katherine Kim
/author/Cassimatis,+Thomas+MThomas M. Cassimatis
A-TITLE Thomas M. Cassimatis
/author/Israni,+Nikita+SNikita S. Israni
A-TITLE Nikita S. Israni
/author/Lea,+Hannah+JHannah J. Lea
A-TITLE Hannah J. Lea
/author/Lentz,+Taylor+NTaylor N. Lentz
A-TITLE Taylor N. Lentz
/author/Pierce,+Anne+EAnne E. Pierce
A-TITLE Anne E. Pierce
/author/Jiang,+AlexAlex Jiang
A-TITLE Alex Jiang
/author/LaMunion,+Samuel+RSamuel R. LaMunion
A-TITLE Samuel R. LaMunion
/author/Thomas,+Reed+JReed J. Thomas
A-TITLE Reed J. Thomas
/author/Ishihara,+AsukaAsuka Ishihara
A-TITLE Asuka Ishihara
/author/Courville,+Amber+BAmber B. Courville
A-TITLE Amber B. Courville
/author/Yang,+Shanna+BShanna B. Yang
A-TITLE Shanna B. Yang
/author/Reitman,+Marc+LMarc L. Reitman
A-TITLE Marc L. Reitman
/author/Cypess,+Aaron+MAaron M. Cypess
A-TITLE Aaron M. Cypess
/author/Chen,+Kong+YKong Y. Chen
A-TITLE Kong Y. Chen
/topic/type/research-articleText duplicate Research Article
/doi/10.1073/pnas.2321179121Effect of skull morphology on fox snow diving
A-TITLE Effect of skull morphology on fox snow diving
/author/Yuk,+JisooJisoo Yuk
A-TITLE Jisoo Yuk
/author/Pandey,+AnupamAnupam Pandey
A-TITLE Anupam Pandey
/author/Park,+LeenaLeena Park
A-TITLE Leena Park
/author/Bemis,+William+EWilliam E. Bemis
A-TITLE William E. Bemis
/author/Jung,+SunghwanSunghwan Jung
A-TITLE Sunghwan Jung
/topic/type/research-articleText duplicate Research Article
/doi/10.1073/pnas.2313797121Atmosphere teleconnections from abatement of China aerosol emissions exacerbate Northeast Pacific warm blob events
A-TITLE Atmosphere teleconnections from abatement of China aerosol emissions exacerbate Northeast Pacific warm blob events
/author/Wang,+HaiHai Wang
A-TITLE Hai Wang
/author/Zheng,+Xiao-TongXiao-Tong Zheng
A-TITLE Xiao-Tong Zheng
/author/Cai,+WenjuWenju Cai
A-TITLE Wenju Cai
/author/Han,+Zi-WenZi-Wen Han
A-TITLE Zi-Wen Han
/author/Xie,+Shang-PingShang-Ping Xie
A-TITLE Shang-Ping Xie
/author/Kang,+Sarah+MSarah M. Kang
A-TITLE Sarah M. Kang
/author/Geng,+Yu-FanYu-Fan Geng
A-TITLE Yu-Fan Geng
/author/Liu,+FukaiFukai Liu
A-TITLE Fukai Liu
/author/Wang,+Chuan-YangChuan-Yang Wang
A-TITLE Chuan-Yang Wang
/author/Wu,+YueYue Wu
A-TITLE Yue Wu
/author/Xiang,+BaoqiangBaoqiang Xiang
A-TITLE Baoqiang Xiang
/author/Zhou,+LeiLei Zhou
A-TITLE Lei Zhou
/topic/type/qnasQnAs
/doi/10.1073/pnas.2412426121QnAs with Spyros Artavanis-Tsakonas
A-TITLE QnAs with Spyros Artavanis-Tsakonas
/author/Ravindran,+SandeepSandeep Ravindran
A-TITLE Sandeep Ravindran
/about/pnas-in-the-newsView all news
/topic/type/brief-reportBrief Report
/doi/10.1073/pnas.2409983121Liquid water in the Martian mid-crust
A-TITLE Liquid water in the Martian mid-crust
/author/Wright,+VashanVashan Wright
A-TITLE Vashan Wright
/author/Morzfeld,+MatthiasMatthias Morzfeld
A-TITLE Matthias Morzfeld
/author/Manga,+MichaelMichael Manga
A-TITLE Michael Manga
/topic/type/research-articleText duplicate Research Article
/doi/10.1073/pnas.2400420121A therapeutic small molecule enhances γ-oscillations and improves cognition/memory in Alzheimer’s disease model mice
A-TITLE A therapeutic small molecule enhances γ-oscillations and improves cognition/memory in Alzheimer’s disease model mice
/author/Wei,+XiaofeiXiaofei Wei
A-TITLE Xiaofei Wei
/author/Campagna,+Jesus+JJesus J. Campagna
A-TITLE Jesus J. Campagna
/author/Jagodzinska,+BarbaraBarbara Jagodzinska
A-TITLE Barbara Jagodzinska
/author/Wi,+DongwookDongwook Wi
A-TITLE Dongwook Wi
/author/Cohn,+WhitakerWhitaker Cohn
A-TITLE Whitaker Cohn
/author/Lee,+Jessica+TJessica T. Lee
A-TITLE Jessica T. Lee
/author/Zhu,+ChunniChunni Zhu
A-TITLE Chunni Zhu
/author/Huang,+Christine+SChristine S. Huang
A-TITLE Christine S. Huang
/author/Molnár,+LászlóLászló Molnár
A-TITLE László Molnár
/author/Houser,+Carolyn+RCarolyn R. Houser
A-TITLE Carolyn R. Houser
/author/John,+VargheseVarghese John
A-TITLE Varghese John
/author/Mody,+IstvanIstvan Mody
A-TITLE Istvan Mody
/topic/type/research-articleText duplicate Research Article
/doi/10.1073/pnas.1321664111Bodily maps of emotions
A-TITLE Bodily maps of emotions
/author/Nummenmaa,+LauriLauri Nummenmaa
A-TITLE Lauri Nummenmaa
/author/Glerean,+EnricoEnrico Glerean
A-TITLE Enrico Glerean
/author/Hari,+RiittaRiitta Hari
A-TITLE Riitta Hari
/author/Hietanen,+Jari+KJari K. Hietanen
A-TITLE Jari K. Hietanen
https://www.pnas.org/trendingView all
/doi/10.1073/pc.32419002play How redlining affects biodiversity
IMG-ALT placeholder
/doi/10.1073/pc.32113667play Animal's eye view of the ocean
IMG-ALT placeholder
/doi/10.1073/pc.31930007play Inequitable exposure to wildfire smoke
IMG-ALT placeholder
/about/science-sessions-podcastview all episodes
/topic/type/opinionOpinion
/doi/10.1073/pnas.2403489121Extended time, elevated expectations: The unappreciated downsides of pausing the tenure clock
A-TITLE Extended time, elevated expectations: The unappreciated downsides of pausing the tenure clock
/author/Holland,+Maria+AMaria A. Holland
A-TITLE Maria A. Holland
/author/Maisel,+KatharinaKatharina Maisel
A-TITLE Katharina Maisel
/author/Ibberson,+Carolyn+BCarolyn B. Ibberson
A-TITLE Carolyn B. Ibberson
/author/Wiley,+Laura+KLaura K. Wiley
A-TITLE Laura K. Wiley
/author/Burnett,+David+CDavid C. Burnett
A-TITLE David C. Burnett
/author/Mace,+Emily+MEmily M. Mace
A-TITLE Emily M. Mace
/author/Elting,+Mary+WilliardMary Williard Elting
A-TITLE Mary Williard Elting
/topic/type/core-conCore Concepts
/doi/10.1073/pnas.2414310121Synthetic data can aid the analysis of clinical outcomes: How much can it be trusted?
A-TITLE Synthetic data can aid the analysis of clinical outcomes: How much can it be trusted?
/author/Adam,+DavidDavid Adam
A-TITLE David Adam
/topic/type/core-conText duplicate Core Concepts
/doi/10.1073/pnas.2413856121Tiny and contentious “crystal clocks” in volcanic rocks could aid early warning for eruptions
A-TITLE Tiny and contentious “crystal clocks” in volcanic rocks could aid early warning for eruptions
/author/McDermott,+AmyAmy McDermott
A-TITLE Amy McDermott
/topic/type/opinionText duplicate Opinion
/doi/10.1073/pnas.2316959121Researchers need to better address time-of-day as a critical biological variable
A-TITLE Researchers need to better address time-of-day as a critical biological variable
/author/Nelson,+Randy+JRandy J. Nelson
A-TITLE Randy J. Nelson
/author/DeVries,+A+CourtneyA. Courtney DeVries
A-TITLE A. Courtney DeVries
/author/Prendergast,+Brian+JBrian J. Prendergast
A-TITLE Brian J. Prendergast
/topic/type/opinionText duplicate Opinion
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Proceedings of the National Academy of Sciences (PNAS), a peer reviewed journal of the National Academy of Sciences (NAS) - an authoritative source of high-impact, original research that broadly spans the biological, physical, and social sciences.

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